1. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture; Wuhan Botanical Garden of Chinese Academy of Sciences; Wuhan 430074 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. Sino-African Joint Research Center; Chinese Academy of Sciences; Wuhan 430074 China
4. Hubei Collaborative Innovation Center for Grain Industry; Jingzhou 434025 China