1. Laboratoire des Ecosystèmes Pastoraux et Valorisation des Plantes Spontanées et des Micro‐organismes Associés Institut des Régions Arides Medenine Tunisia
2. Department of Biology, Faculty of Sciences University of Gabes Gabès Tunisia
3. Food Engineering Division National Food Research Institute, NARO Tsukuba Japan
4. ARENA (Alliance for Research for North Africa) Tsukuba University Tsukuba Japan