1. Department of Plant Reproductive Biology and Epigenetics Max Planck Institute of Molecular Plant Physiology Potsdam 14476 Germany
2. Department of Plant Biotechnology and Bioinformatics Ghent University Ghent 9052 Belgium
3. VIB Center for Plant Systems Biology, VIB Ghent 9052 Belgium
4. Department of Plant Biology, Uppsala BioCenter Swedish University of Agricultural Sciences and Linnean Centre for Plant Biology Uppsala 75007 Sweden