1. Laboratory for Integrated Cellular Systems; RIKEN Center for Integrative Medical Sciences (IMS-RCAI); Tsurumi-ku Yokohama Kanagawa 230-0045 Japan
2. Division of Cell Proliferation, ART; Graduate School of Medicine; Tohoku University; Sendai Miyagi 980-8575 Japan
3. Department of Surgery; University of Michigan; Ann Arbor MI 48109 USA
4. Department of Microbiology and Immunology; University of Michigan; Ann Arbor MI 48109 USA
5. Laboratory for Lymphocyte Differentiation; RIKEN Center for Integrative Medical Sciences (IMS-RCAI); Tsurumi-ku Yokohama Kanagawa 230-0045 Japan
6. Laboratory for Lymphocyte Differentiation; Immunology Frontier Research Center; Osaka University; 3-1 Yamada-oka Suita Osaka 565-0871 Japan