1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany Chinese Academy of Sciences Kunming Yunnan 650201 China
2. Department of Economic Plants and Biotechnology Yunnan Key Laboratory for Wild Plant Resources Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
3. Global Biodiversity Conservancy 144/124 Moo3, Soi Bua Thong Bangsalae, Sattahip Chonburi 20250 Thailand
4. Institute of Ecology and Key Laboratory for Earth Surface Processes of the Ministry of Education College of Urban and Environmental Sciences Peking University Beijing 100871 China
5. Movement Ecology Laboratory Department of Ecology, Evolution and Behavior Alexander Silberman Institute of Life Sciences The Hebrew University of Jerusalem Jerusalem 91904 Israel
6. University of Chinese Academy of Sciences Beijing 100049 China