1. Institute for the Environment; Brunel University London; Uxbridge UB8 3PH UK
2. Department of Environmental Sciences (IMV); Norwegian University of Life Sciences (NMBU); NO-1432 Ås Norway
3. Department of Ecology and Natural Resource Management (INA); Norwegian University of Life Sciences (NMBU); NO-1432 Ås Norway
4. Experimental Techniques Centre; Brunel University London; Uxbridge UB8 3PH UK