1. Department of Zoology; Graduate School of Science; Kyoto University; Kyoto Japan
2. Graduate School of Science and Technology; Shizuoka University; Hamamatsu Japan
3. Department of Environmental and Forest Biology; College of Environmental Science and Forestry; State University of New York; Syracuse NY USA
4. Marine Biosystems Research Center; Chiba University; Uchiura Kamogawa Chiba Japan
5. Department of Ecology and Evolutionary Biology; University of Connecticut; Storrs CT USA