1. Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut 06269, USA;
2. Department of Ecology and Evolutionary Biology, University of Connecticut, Hartford, Connecticut 06103, USA;
3. Department of Entomology and Nematology, University of California, Davis, California 95616, USA;
4. Department of Zoology, Graduate School of Science, Kyoto University, Sakyo, Kyoto 606-8502, Japan;