1. University of Groningen; Department of Ocean Ecosystems; PO Box 11103, 9700 CC Groningen The Netherlands
2. The Swire Institute of Marine Science and School of Biological Sciences; The University of Hong Kong; Cape d'Aguilar Shek O Hong Kong
3. Nelson Mandela Metropolitan University; Coastal and Marine Research Institute; PO Box 77000 Port Elizabeth 6031 South Africa
4. South African Institute for Aquatic Biodiversity (SAIAB) and South African Environmental Observation Network (SAEON); Grahamstown 6140 South Africa