1. Department of Molecular and Cell Biology, University of Cape Town, Rondebosch 7700, South Africa
2. British Antarctic Survey, Cambridge CB3 0ET, UK
3. Natural Science Collections Facility (South African National Biodiversity Institute), Pretoria 0186, South Africa
4. Department of Natural History, Iziko South African Museums, Cape Town 8001, South Africa
5. Bayworld Centre for Research and Education (BCRE), Port Elizabeth 6013, South Africa
6. Division of Medical Virology, Department of Pathology, Stellenbosch University, Cape Town 7505, South Africa