1. Halpin Wildlife Research; Vancouver British Columbia V6G 1J3 Canada
2. Department of Biology; Dalhousie University; Halifax Nova Scotia B3H 4R2 Canada
3. Department of Botany and Biodiversity Research Centre; University of British Columbia; Vancouver British Columbia V6T 1Z4 Canada
4. Canadian Wildlife Service; Environment and Climate Change Canada; Sidney British Columbia V8L 4B2 Canada
5. Carter Biological Consulting; Victoria British Columbia V8S 4S8 Canada