1. School of Natural & Environmental Sciences, Ridley Building; Newcastle University; Newcastle upon Tyne NE1 7RU U.K.
2. Environmental Research Unit; University of Highlands and Islands; Thurso KW14 7EE U.K.
3. Institute for Ageing, Campus for Ageing and Vitality; Newcastle University; Newcastle upon Tyne NE4 5PL U.K.
4. Center for Healthy Aging, Department of Cellular and Molecular Medicine; University of Copenhagen; Copenhagen 2200 Denmark