1. Department of Fisheries Faculty of Marine Natural Resources Khorramshahr University of Marine Science and Technology Khorramshahr Iran
2. South Iran Aquaculture Research Centre Iranian Fisheries Science Institute (IFSRI) Agricultural Research Education and Extension Organization (AREEO) Ahwaz Iran
3. Department of Basic Sciences Faculty of Veterinary Medicine Shahid Chamran University of Ahvaz Ahvaz Iran