1. Department of Biosciences; University of Eastern Finland; Kuopio Finland
2. Department of Biological and Environmental Science; University of Jyväskylä; Jyväskylä Finland
3. Department of Environmental Science; University of Eastern Finland; Kuopio Finland
4. Finnish Game and Fisheries Research Institute; Jyväskylä Game and Fisheries Research; Jyväskylä Finland