1. Division of Genetics and Physiology; Department of Biology; University of Turku; Itäinen Pitkäkatu 4 Turku FIN-20520 Finland
2. Finnish Game and Fisheries Research Institute; Utsjoki FIN-99980 Finland
3. Centre for Integrative Genetics (CIGENE) and Department of Animal and Aquacultural Sciences; Norwegian University of Life Sciences; Aas N-1432 Norway