1. Key Laboratory of Ecosystem Network Observation and Modeling; Institute of Geographic Sciences and Natural Resources; the Chinese Academy of Sciences; Beijing China
2. Department of Ecology, Evolution and Natural Resources; Rutgers University; New Brunswick NJ 08901 USA
3. University of Chinese Academy of Sciences; Beijing China
4. Key Laboratory of the Zoological Systematics and Evolution; Institute of Zoology; the Chinese Academy of Sciences; Beijing China
5. PRC-GEF Partnership on Land Degradation in Dryland Ecosystems; Beijing China