1. School of Planning, Architecture and Civil Engineering; Queen's University Marine Laboratory; 12-13 The Strand Portaferry BT22 1PF UK
2. Department of Marine Science; University of Otago; PO Box 56 Dunedin 9054 New Zealand
3. School of Biological Sciences; Queen's University Marine Laboratory; 12-13 The Strand Portaferry BT22 1PF UK