1. Environment and Sustainability Institute; University of Exeter; Penryn Campus Penryn TR10 9FE Cornwall UK
2. Department of Biology; University of York; York YO10 5DD UK
3. Department of Geography; University of Exeter; Penryn Campus Penryn TR10 9FE Cornwall UK
4. Centre for Ecology and Conservation; University of Exeter; Penryn Campus Penryn TR10 9FE Cornwall UK
5. People's Trust for Endangered Species; 3 Cloisters House, 8 Battersea Park Road, Battersea London SW8 4BG UK
6. Forest Enterprise Scotland; Dumfries and Borders Forest District; Ae Village Parkgate Dumfries DG1 1QB UK