1. Crop Research Institute Shandong Academy of Agricultural Sciences Jinan China
2. Ministry of Agriculture Key Laboratory of Wheat Biology and Genetic Improvement on North Yellow and Huai River Valley Jinan China
3. National Engineering Laboratory for Wheat and Maize Jinan Shandong China
4. Department of Plant Science and Landscape Architecture University of Maryland College Park MD USA
5. Institute for Bioscience and Biotechnology Research University of Maryland Rockville MD USA