1. Department of Genetics, Cell Biology, and Development and Center for Genome Engineering, University of Minnesota, Minneapolis, Minnesota 55455
2. Department of Plant Pathology, University of Minnesota, St. Paul, Minnesota 55108
3. Department of Agronomy and Plant Genetics, University of Minnesota, St. Paul, Minnesota 55108
4. Department of Plant Developmental Genetics, Institute of Biophysics of the CAS, CZ-61265 Brno, Czech Republic