1. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture; Wuhan Botanical Garden of the Chinese Academy of Sciences; Wuhan 430074 China
2. Graduate University of Chinese Academy of Sciences; 19A Yuquanlu Beijing 100049 China
3. The New Zealand Institute for Plant and Food Research Ltd, (Plant and Food Research); Mt Albert Research Centre; Private Bag 92169 Auckland New Zealand
4. Institute of Fruit Tree and Tea; Hubei Academy of Agricultural Sciences; 430209 Wuhan China