1. Key Laboratory of Bio‐Resource and Eco‐Environment of Ministry of Education College of Life Sciences State Key Laboratory of Hydraulics and Mountain River Engineering Sichuan University Chengdu Sichuan 610065 China
2. Royal Botanic Garden Edinburgh 20A Inverleith Row Edinburgh EH3 5LR UK
3. Institute of Molecular Plant Sciences The University of Edinburgh Edinburgh EH9 3JH UK
4. College of Science Tibet University Lhasa Xizang Autonomous Region 850012 China