1. Centre d'Estudis Avançats de Blanes (CEAB-CSIC); Accés a la Cala S. Francesc 14 17300 Blanes Girona Spain
2. Ocean and Earth Science; University of Southampton; National Oceanography Centre; Southampton European Way, Southampton SO14 3ZH UK
3. Molecular Ecology Laboratory; School of Biological Sciences; Flinders University; Adelaide SA 5001 Australia