1. Center of Forest Ecosystem Studies and Qianyanzhou Ecological Station Key Laboratory of Ecosystem Network Observation and Modeling Synthesis Research Center of Chinese Ecosystem Research Network Institute of Geographic Sciences and Natural Resources Research Chinese Academy of Sciences Beijing 100101 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Department of Ecology College of Urban and Environmental Sciences, and Key Laboratory for Earth Surface Processes of the Ministry of Education Peking University Beijing 100871 China
4. Department of Biology Brigham Young University Provo UT 84602 USA
5. Department of Ecosystem Science and Management, and Intercollege Graduate Degree Program in Plant Biology Pennsylvania State University University Park PA 16802 USA