1. Department of Plant Sciences, Laboratory of Molecular Biology, Wageningen University, Droevendaalsesteeg 1, 6708 PB Wageningen, Netherlands.
2. College of Science, King Saud University, Post Office Box 2455, Riyadh 11451, Saudi Arabia.
3. Plant Developmental Biology, Wageningen University, Droevendaalsesteeg 1, 6708 PB Wageningen, Netherlands.