1. Key Laboratory of Experimental Marine Biology; Institute of Oceanology; Chinese Academy of Sciences; Qingdao 266071 China
2. Laboratory for Marine Biology and Biotechnology; Qingdao National Laboratory for Marine Science and Technology; Qingdao 266237 China
3. University of Chinese Academy of Sciences; Beijing 100049 China
4. Department of Biological Sciences; University of Texas at Dallas; Dallas Texas 75080 USA