1. LEMAR UMR 6539 UBO CNRS IRD; European Institute of Marine Studies (IUEM); University of Western Brittany (UBO); European University of Brittany (UEB); Technopôle Brest-Iroise Plouzané 29280 France
2. RMN-RPE-SM; 6 avenue, Victor-Le-Gorgeu-CS93837 Brest Cedex 3 29238 France
3. UMR 7138 Systématique, Adaptation, Evolution; IRD; B.P. A5 Nouméa Cedex 98848 Nouvelle-Calédonie
4. UPF; French Polynesia