1. Department of Biology University of Toronto Mississauga 3359 Mississauga Road Mississauga Ontario L5L 1C6 Canada
2. Department of Cell & Systems Biology University of Toronto 25 Harbord Street Toronto Ontario M5S 3G5 Canada
3. California Institute for Quantitative Biosciences University of California Berkeley, 174 Stanley Hall Berkeley California 94720‐3220 USA
4. Department of Ecology and Evolutionary Biology University of Toronto 25 Willcocks Street Toronto Ontario M5S 3B2 Canada