1. Department of Marine Systems, Tallinn University of Technology, Tallinn 12618, Estonia
2. The Royal Society for the Protection of Birds, 2 Lochside View, Edinburgh EH12 9DH, Scotland, UK
3. Aarhus Institute of Advanced Studies, Aarhus University, Building 1630, Aarhus 8000, Denmark
4. Department of Biology, Aarhus University, Building 1135, Aarhus 8000, Denmark