1. Department of Zoology; Graduate School of Science; Kyoto University; Kitashirakawa Sakyo-ku Kyoto 606-8502 Japan
2. Graduate School of Biological Sciences; Nara Institute of Science and Technology; Takayama Ikoma Nara 630-0192
3. Core Research for Evolutional Science and Technology (CREST); Japan Science and Technology Agency (JST); Chiyoda-ku Tokyo 102-0076 Japan