1. State Key Laboratory for Agrobiotechnology; Key Laboratory of Crop Heterosis and Utilization (MOE); Beijing Key Laboratory of Crop Genetic Improvement; China Agricultural University; Beijing 100193 China
2. National Plant Gene Research Centre (Beijing); Beijing 100193 China
3. Department of Plant Genetics & Breeding; China Agricultural University; Yuanmingyuan Xi Road No. 2; Haidian District, Beijing 100193 China
4. Institute of Plant Science Paris-Saclay; Université Paris Sud; 91405 Orsay France
5. National Maize Improvement Centre of China; China Agricultural University; Beijing 100193 China