1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla 666303 Yunnan China
2. Center of Plant Ecology, Core Botanical Gardens, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla 666303 Yunnan China
3. Germplasm Bank of Wild Species, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
4. Department of Botany Smithsonian Institution Washington DC 20013 USA
5. Department of Anthropology University of Toronto Scarborough Toronto M5H 2N2 Canada
6. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650204 China