1. Biodiversity Research Group; School of Geography and the Environment; Oxford University Centre for the Environment; University of Oxford; South Parks Road Oxford OX1 3QY U.K.
2. G.B. Pant Institute of Himalayan Environment and Development; Kosi-Katarmal Almora 263643 Uttarakhand India
3. Plant Diversity; Systematics and Herbarium Division; CSIR-National Botanical Research Institute; Rana Pratap Marg Lucknow 226001 India
4. Center for Macroecology Evolution and Climate; University of Copenhagen; Universitetsparken 15; DK-2100 Copenhagen Denmark