1. CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture Wuhan Botanical Garden Chinese Academy of Sciences Wuhan Hubei 430074 China
2. Center of Conservation Biology Core Botanical Gardens Chinese Academy of Sciences Wuhan Hubei 430074 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Department of Botany and Plant Sciences University of California Riverside Riverside CA 92507 USA
5. School of Integrative Plant Science Section of Plant Biology and the L.H. Bailey Hortorium Cornell University Ithaca NY 14850 USA
6. Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany Chinese Academy of Sciences Kunming Yunnan 650201 China
7. Hubei Key Laboratory of Shennongjia Golden Monkey Conservation Biology Administration of Shennongjia National Park Shennongjia Hubei 442400 China