1. Behaviour and Evolution, Ethology Lab, Cavanilles Institute of Biodiversity and Evolutionary Biology University of Valencia C/Catedrático José Beltrán 2 Paterna Valencia 46980 Spain
2. Doñana Biological Station Spanish Research Council CSIC c/Americo Vespucio, 26, Isla de la Cartuja Sevilla 41092 Spain
3. Centre for Evolutionary Biology, School of Biological Sciences The University of Western Australia 35 Stirling Highway Crawley Western Australia 6009 Australia
4. Ecology and Evolution Research Centre, School of Biological, Earth and Environmental Sciences University of New South Wales Sydney New South Wales 2052 Australia
5. Division of Ecology and Evolution, Research School of Biology The Australian National University Canberra Australian Capital Territory 2061 Australia