1. Center for Integrative Conservation Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Mengla 666303 Yunnan China
2. Center of Conservation Biology, Core Botanical Gardens Chinese Academy of Sciences Mengla 666303 Yunnan China
3. Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences Yezin Nay Pyi Taw 05282 Myanmar
4. Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences Guilin 541006 Guangxi China
5. Germplasm Bank of Wild Species Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
6. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
7. Department of Biological Sciences Sam Houston State University Huntsville TX 77341 USA