1. Department of Plant Pathology; Faculty of AgriSciences; Stellenbosch University; Private Bag X1 Matieland 7602 South Africa
2. Evolution, Ecology and Genetics; Research School of Biology; The Australian National University; 116 Daley Road Canberra ACT 2601 Australia
3. Citrus Research International; PO Box 28 Nelspruit 1200 South Africa