1. The Sainsbury Laboratory University of Cambridge Bateman Street Cambridge CB1 2LR UK
2. Department of Biology University of Massachusetts 611 North Pleasant Street Amherst MA 01003‐9297 USA
3. Department of Molecular, Cell and Developmental Biology University of California at Los Angeles 610 Charles E Young Dr. South Los Angeles CA 90095 USA
4. Molecular Biology Institute University of California at Los Angeles 611 Charles E. Young Drive East Los Angeles CA 90095‐1570 USA