1. School of GeoSciences; University of Edinburgh; Drummond Street; Edinburgh; EH8 9XP; Scotland; UK
2. Estonian University of Life Sciences; 51041; Tartu; Estonia
3. Alterra, Wageningen University and Research Centre; PO Box 47; 6700 AA; Wageningen; The Netherlands
4. Climate and Land Use Change Program; United States Geological Survey; 12201 Sunrise Valley Drive; MS 519; Reston; VA; 20192; USA
5. Centre for Mountain Ecosystems Studies; Kunming Insitute of Botany; China