1. Department of Zoology University of Cambridge CambridgeCB2 3EJUK
2. Research Centre of Ecological Change Faculty of Biological and Environmental Sciences University of Helsinki Helsinki Finland
3. Helsinki Institute of Life Science HiLIFEUniversity of Helsinki Helsinki Finland
4. Department of Biological and Medical Sciences Oxford Brookes University OxfordOX3 0BPUK
5. School of Health and Life Sciences University of West Scotland PaisleyPA1 2BEScotland