1. Department of Biology; Stanford University; Herrin Labs 454 Stanford CA 94305 USA
2. Université de Lyon F-69000 Lyon; Université Lyon 1; CNRS, UMR5558; Laboratoire de Biométrie et Biologie Évolutive; F-69622 Villeurbanne France
3. Department of Zoology; University of Oxford; Tinbergen Building South Parks Road Oxford OX1 3PS UK