1. Departamento de Biología de Organismos y Sistemas Universidad de Oviedo Oviedo Spain
2. Research Unit of Biodiversity (UO‐CSIC‐PA)Campus de Mieres Mieres Spain
3. School of Biological Sciences Monash University Clayton Vic. Australia
4. School of BioSciences The University of Melbourne Parkville Vic. Australia
5. Department of Evolutionary Ecology Estación Biológica de DoñanaCSIC Sevilla Spain