1. Centre for Ecology and Conservation; Penryn Campus; University of Exeter; Penryn TR10 9EZ UK
2. Conservation Biology Division; Northwest Fisheries Science Center; National Marine Fisheries Service; National Oceanic and Atmospheric Administration; 2725 Montlake Boulevard East Seattle WA 98112 USA
3. British Antarctic Survey; Natural Environment Research Council; High Cross, Madingley Road Cambridge CB3 0ET UK
4. SUERC; Rankine Avenue, Scottish Enterprise Technology Park East Kilbride G75 0QF UK