1. Lancaster Environment Centre; Lancaster University; Lancaster UK
2. Department of Zoology; Edward Grey Institute; University of Oxford; Oxford UK
3. CNRS, UMR 5558 “Biométrie et Biologie Evolutive”; Université de Lyon; Villeurbanne France
4. Rubenstein School of Environment and Natural Resources; University of Vermont; Burlington VT USA
5. British Ecological Society; London UK