1. CNRS UMR 5558 Laboratoire de Biométrie et Biologie Evolutive Université de LyonUniversité Lyon 1 Villeurbanne France
2. Rubenstein School of Environment and Natural Resources University of Vermont Burlington VT USA
3. Institute of Biological and Environmental Sciences The University of Aberdeen Aberdeen UK
4. School of Natural and Environmental Sciences Newcastle University Newcastle upon Tyne UK
5. British Ecological Society London UK