1. Kunpeng Institute of Modern Agriculture at Foshan Foshan 528200 Guangdong China
2. Shenzhen Branch, Guangdong Laboratory for Lingnan Modern Agriculture, Genome Analysis Laboratory of the Ministry of Agriculture Agricultural Genomics Institute at Shenzhen Shenzhen 518120 Guangdong China
3. Department of Agricultural Biology Colorado State University Fort Collins CO 80523 USA
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing 100093 China