1. Department of Plant and Soil Sciences University of Pretoria Pretoria South Africa
2. Institute for Biodiversity and Ecosystem Dynamics University of Amsterdam Amsterdam The Netherlands
3. School of Animal, Plant and Environmental Sciences University of the Witwatersrand Johannesburg South Africa
4. FABI (Forestry and Agricultural Biotechnology Institute) University of Pretoria Pretoria South Africa
5. Centre for Biodiversity Dynamics in a Changing World Aarhus University Aarhus C Denmark