1. Plant Pathology, Institute of Integrative Biology, ETH Zurich; 8092 Zurich Switzerland
2. Syngenta Crop Protection AG; Schaffhauserstrasse 4332 Stein Switzerland
3. School of Molecular Bioscience; University of Sydney; Sydney NSW Australia
4. Department of Plant Biology and Crop Science; Rothamsted Research; Harpenden Hertfordshire AL5 2JQ UK