1. State Key Laboratory of Plateau Ecology and Agriculture Qinghai University Xining PR China
2. Ministry of Education Key Laboratory of Western China's Environmental Systems Lanzhou University Lanzhou PR China
3. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering School of Life Sciences, Fudan University Shanghai PR China
4. Key Laboratory of Adaptation and Evolution of Plateau Biota Northwest Institute of Plateau Biology, Chinese Academy of Sciences Xining PR China