1. Fish Ecology and Conservation Physiology Laboratory, Department of Biology and Institute of Environmental Science; Carleton University, 1125 Colonel By Drive; Ottawa ON K1S 5B6 Canada
2. Danish Centre for Wild Salmon, Brusgårdsvej 15; 8960 Randers Denmark
3. DTU AQUA, National Institute of Aquatic Resources, Section for Freshwater Fisheries and Ecology; Technical University of Denmark, Vejlsøvej 39; 8600 Silkeborg Denmark
4. Department of Biology; University of Ottawa, 30 Marie-Curie; Ottawa ON K1N 6N5 Canada